From Kirk McKusick:

  If a directory somehow develops a hole (that is a block pointer
  that has a value of zero), fsck would give the filesystem a clean
  bill of health, but the kernel would panic when accessing the
  directory with the hole. Fsck now checks for holes in directories.
  If found in preen mode, fsck fails. In manual mode, it can be
  directed to shorten the directory to the beginning of the hole. A
  more complete solution would be to allocate a block to fill the
  hole. However, this is a lot more work for a `cannot happen' error,
  so the extra effort seems unwarranted.

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